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Krautter, M. (2002): Fossil Hexactinellida: An
Overview. In: Hooper,
J.N.A. & Van Soest, R.W.M. (eds): Systema Porifera: A Guide to the
Classification
of Sponges, 1211-1223; Kluwer Academic/Plenum Publishers (New York).
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Abstract: Hexactinellida (Porifera)
are known from hexactin spicules from the Late Proterozoic
of Mongolia (Brasier et al., 1997) and China (Steiner et al., 1993), possibly
representing the oldest lineage of animals alive on earth today.
Hexactinellida flourished and radiated rapidly during the Middle Cambrian times,
giving rise to many new taxa and new skeletal plans (Rigby, 1986c; Walcott, 1920).
Our knowledge of Early Paleozoic Hexactinellida is largely derived from isolated
spicules from sediments. Both Amphidiscophora and Hexasterophora are known since the
Early Paleozoic but most Paleozoic families did not survive the Paleozoic-Mesozoic
boundary. Lyssacinosida are the first Hexactinellida known since the Proterozoic. They
are an ultra-conservative group and their body-plan still exists in living species today.
The first representatives of Hexactinosida are known since the Devonian, with maximum radiation
and diversity occurring during the Mesozoic. The first lychniscosidan sponges
were described from the Middle Jurassic of Europe and during the Cretaceous
where they reached their maximum peak of diversity. After the Jurassic-Cretaceous
there has been a gradual decline in diversity leading up to the present. The present
overview discusses about 120 genera in 18 families and two orders.
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