In the Lusitanian Basin (south of Coimbra), the marl-limestone alternations
of the middle Toarcian/lower Aalenian show the occurrence of small-
-scale siliceous sponge mud mounds.
These facies are particularly well documented and preserved, belonging
to the following two units: Marls and marly limestones with sponge reefs
(S. Gião Fm) and to the lower part of the Póvoa da Lomba Fm. Both units
correspond to very bioturbated (Chondrites, Zoophycos, Planolites and
Thalassinoides) marl-limestone successions. Between these two units
exists the Marls and marly limestone with brachiopods (top of S. Gião
Fm) wich corresponds to a marly unit, with rare limestone levels and
are very poor in siliceous sponge reefs. This formation is dated of
the uppermost Bifrons Zone through the base of the Bonarellii Zone.
The majority of the siliceous sponge mud mounds occur within this time
slice. These mounds are characterized by a great abundance and diversity
of benthic macrofauna mainly composed of brachiopods (rhynchonellids
and terebratulids), crinoids and bivalves. The base (Meneghini-Opalinum
interval) of the Póvoa da Lomba Fm is composed by bioturbated marl/limestone
alternations, with an upward increase of calcareous (biomicrite/wackestone,
locally packstone) facies. The rhythmicity of the marl/limestone couplets
is an important feature, defining elementary sequences (parasequences).
The vertical succession of the parasequences is organised into parasequence
groups showing a clear asymmetry, with the marls typically thicker at
the base. Besides, the tops of these fourth-order sequences are marked
by a thickening of the limestone beds and, sometimes, by a large density
of Zoophycos. The siliceous sponge mud mound occurrences are particularly
associated with these levels.
| 
Map of Portugal.
The siliceous sponge mud mounds
are located just south of Coimbra
(asterisk). |

Litho- and biostratigraphy of the Toarcian-
Lower Aalenian from the Lusitanian Basin. |
The Toarcian mud mounds of the Lusitanian Basin are usually only a
few decimetres thick and most display irregular knob -like to flat lenticular
morphologies. Some build-ups are round and can reach 1.5 metres in thickness
and ten metres in diameter. The upper mound surface is normally rough
and uneven. In both sequences they are always related laterally with
carbonate beds, which correspond preferentially to the tops of fourth-order
sequences. The mud mounds consist of mostly brownish, iron-rich calcified
siliceous sponges and a greyish, sometimes peloidal allochthonous micritic
matrix. In general, the sponges themselves consist of dense leiolitic
microbolites (automicrites sensu Reitner & Neuweiler, 1993). The
sponge spicules are diagenetically transformed into calcite. The great
majority of the sponge specimens belongs to the Hexactinosida (Class
Hexactinellida) and are unknown and undescribed to date. “Lithistides”
(polyphyletic desma-bearing demosponges) are very rare and only occur
as forms encrusting hexactinosidan sponges. The benthic macrofauna is
abundant and consists of monospecific crinoids, rhynchonellids, terebratulids
and bivalves (mainly pectinids and ostreids). Encrusting organisms are
serpulids, bryozoans and foraminifera, as well as “Lithistids”,
mentioned above. They are entirely restricted to the stratal surfaces
of the siliceous sponges. The sponge reefs consist of several microfacies
types (wackestones, packstones, floatstones and boundstones). All of
them are micrite dominated and represent low energy environments. They
differ mainly in the amount of siliceous sponges, micrite, microbialites,
encrusting organisms and the accompanying fauna.
Palaeoenvironmental Significance
The amount of microbially induced carbonate clearly emphasizes the importance
of microbial activity in respect of the reef building potential. Furthermore,
three other controlling factors played an important role in the initiation
of the siliceous sponge mud mounds of the Lusitanian Basin: bathymetry,
sea-floor morphology and sedimentation rate. The role of the first two
factors is evident because the siliceous sponge mud mounds are particularly
important (abundance and volumetric expression) in the eastern part
of the basin (Rabaçal-Alvaiázere region). They are practically
absent towards the west (essentially in the base of Póvoa da
Lomba Fm) where the series shows hemipelagic features. Reduced sedimentation
rate was a precondition for the settlement of siliceous sponges and
Hexactinosida in particular. The preferential occurrence of these bioconstructions
at the top of fourth order sequences seems to be related with sediment-starved
intervals.
Pictures (click the thumbs to enlarge)
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typical small-scaled knob-like sponge mud mound in the Lusitanian
Basin of Portugal |
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small-scaled lentille-shaped sponge mud mound in the Lusitanian
Basin of Portugal |
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small knob-like sponge mud mound in the Lusitanian Basin of Portugal |
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undiscribed hexactinosidan sponge |
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Bioclast-rich sponge floatstone with various sponge specimens
partly stained with iron-oxide |
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"Baby"-sponge reef, formed by several small hexactinosidan
sponges which grow upon each other |
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