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- Phylogeny and evolution of man
- Biostratigraphy and phylogenetic reconstructions
- Peleoecology of marine invertebrates
- Benthic habitat studies
- Sedimentology and diagenesis of limestones
- Evolution of early life
- Crisis in the evolution of life

Research collaboration with Geo- and Bioscience Institutions from Germany,
Switzerland, Austria, Italy, Spain, Portugal, Poland, France, Ireland,
Great Britain, Argentina, Australia, USA and Canada.
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